An ellipse rotating rigidly about its center may appear to rotate rigidly or to deform nonrigidly so that it appears gelatinous. We use this ambiguous stimulus to study how motion information is propagated across space. We find that features that are quite far from the ellipse's contour may have a strong influence on the ellipse's percept, provided they move in a way consistent with the ellipse's motion. We show that the percept cannot be accounted for by computational models that only pool constraints over a local area, nor by models that propagate information along contours nor by models that indiscriminately propagate information across space. However the percept can be accounted for by a class of models that assume smoothness in a layered representation.